On the peer-reviewed level--
R. Dudley, P. Chai, and others performed experiments within the last
decade with hummingbirds and reduced flight-medium density which in my
opinion can only be classified as elegant. Of particular interest is
confirmation of the intuitive perception that wing-stroke amplitude and
frequency increase as flight-medium density decreases. The corollary is
that increased atmospheric density reduces wing-stroke
amplitude/frequency, with obvious implications for take-off scenarios in
large animals with long wings.
Don