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RE: Sereno's (2005) new definitions
Mickey Mortimer wrote:
A nomen dubium cannot influence a topology except to create a polytomy
which covers its potential placements. Only taxa with unique combinations
of characters can influence topologies, but nomina dubia by definition lack
those.
Depends who you ask. :-) I would say that a unique combination of
characters is enough to uphold the validity of a taxon - and I'm sure you
would too. But there is another (more stringent) view that a taxon must
have autapomorphies in order to be deemed valid. That seems a little harsh
to me.
In other words, we could put a nomen dubium into a matrix - but why would
we need to? Why, when we could just replace the nomen dubium with an
unquestionably valid taxon?
To tell what the nomen dubium's phylogenetic relationships are, of course.
But by definition, a nomen dubium does not represent a discrete organism.
Because a nomen dubium has no unique characters (or combination of
characters) it cannot be differentiated from other taxa. For example,
_Antrodemus_ might be _Allosaurus_, or it may not be; we're never going to
know. Therefore it is weird (for want of a better word) to include a nomen
dubium in a matrix. I think this is what Jaime is also driving at: a nomen
dubium has nomenclatural standing (in the sense that the name is valid) but
no taxonomic standing (in the sense that we don't know what organism it
pertains too).
Why view -idae and -ia clades differently if we're ignoring the ICZN?
Because -idae clades should at least be assured of containing the eponymous
genus. This goes way beyond the ICZN: the suffix -idae represents a
tradition going back thousands of years. The ancient Greeks used '-idae' to
designate a dynastic lineage or family: Atreidae, Heraclidae, Achaemenidae,
Seleucidae, Arsacidae, etc. Each lineage/family had the eponymous ruler or
hero as a member - in these cases, this was the founder (real or alleged).
So, to answer your question, -idae and -ia should be viewed differently
because -idae actually denotes something in its own right.
Cheers
Tim