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I agree with Tim. Just expand Sauropodomorpha, since herrerasaurids were sauropodomorphs from right after Herrerasaurus' description until the cladistic revolution anyway.
Pachypodosauria is as bad as Ornithoscelida, since it was originally conceived by Huene as a group combining carnosaurs and sauropodomorphs to the exclusion of coelurosaurs. Thus a Pachypodosauria without carnosaurs doesn't match the concept. Kischlat
(2001) similarly defined Pachypodosauria as everything closer to Morosaurus than Allosaurus. While his thesis (2003) proposed this in more depth and implied Allosaurus was not mentioned by Huene, anyone familiar with historical classification knows that Huene
included Allosaurus in his pachypodosaurian 'Megalosauriden (s. lat.)'. So Allosaurus wasn't explicitly included in Pachypodosauria by Huene, but really it was. Mickey Mortimer From: dinosaur-l-request@usc.edu <dinosaur-l-request@usc.edu> on behalf of Tim Williams <tijawi@gmail.com>
Sent: Thursday, May 4, 2017 6:45 PM To: dinosaur-l@usc.edu Subject: [dinosaur] Ornithoscelida sister clade: Sauropodomorpha or Pachypodosauria, not Saurischia This just out (well done Tom!)...
Holtz, T.R. Nomenclature: Share names for dinosaur divisions. Nature 545: 30 doi:10.1038/545030d First paragraph: "Matthew Baron and colleagues propose a radical revision of dinosaur relationships (Nature 543, 501-506; 2017). I suggest that borrowing from the field's rich history could help to prevent unnecessary confusion." Holtz argues that Saurischia should be abandoned if it doesn't include Theropoda - I completely agree. He further suggests that the name Sauropodomorpha, or alternatively Pachypodosauria, should be used for the remaining 'saurischians' (= traditional sauropodomorphs + more basal taxa, like herrerasaurids). I like the rationale for reviving Huene's Pachypodosauria ("Initially used to unite larger theropods with the sauropodomorphs, the term could now be applied to a clade of dinosaurs that have four or more weight-bearing toes, as distinct from their lighter-footed ornithoscelidan sister taxon with three weight-bearing toes."). But I think the preferred option is to use Sauropodomorpha for this clade, rather than Pachypodosauria. Sauropodomorpha would simply be expanded to include herrerasaurids (or whichever taxa are recovered as closer to sauropods than to theropods or ornithischians). IIRC, Pachypodosauria originally included teratosaurids, many of which were based on the erroneous combination of predatory teeth (including from rauisuchians) and associated basal sauropodomorph ('prosauropod') bones. Hence these teratosaurids were regarded by Huene as primitive carnivores that provided a link between large theropods (Huene's 'carnosaurs') and sauropods. I don't think this historical 'baggage' interferes with resurrecting Pachypodosauria as a new dinosaurian clade. However, given that inclusion of herrerasaurids in the Sauropodomorpha has precedent, I think Sauropodomorpha is the best way to go for the name of the sister taxon to Baron et al.'s Ornithoscelida. |