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A new Liaoning pterosaur: Elanodactylus prolatus gen. et sp. nov.
BRIAN ANDRES, JI QIANG (2008) A NEW PTEROSAUR FROM THE LIAONING PROVINCE OF
CHINA, THE PHYLOGENY OF THE PTERODACTYLOIDEA, AND CONVERGENCE IN THEIR
CERVICAL VERTEBRAE
Palaeontology 51 (2) , 453-469
Don't have this one yet, but here's the abstract:
The largest known flying organisms are the azhdarchid pterosaurs, a
pterodactyloid clade previously diagnosed by the characters of their
extremely elongate middle-series cervical vertebrae. The named species of
the Azhdarchidae are from the Late Cretaceous. However, isolated
mid-cervical vertebrae with similar dimensions and characters have been
referred to this group that date back to the Late Jurassic, implying an
almost 60 million year gap in the fossil record of this group and an
unrecorded radiation in the Jurassic of all the major clades of the
Pterodactyloidea. A new pterosaur from the Early Cretaceous of Liaoning
Province of China, Elanodactylus prolatus gen. et sp. nov., is described
with mid-cervical vertebrae that bear these azhdarchid characters but has
other postcranial material that are distinct from the members of this group.
Phylogenetic analysis of the new species and the Pterodactyloidea places it
with the Late Jurassic vertebrae in the Late Jurassic-Early Cretaceous
Ctenochasmatidae and reveals that the characters of the elongate azhdarchid
vertebrae appeared independently in both groups. These results are realized
though the large taxon sampling in the analysis demonstrating that the
homoplastic character states present in these two taxa were acquired in a
different order in their respective lineages. Some of these homoplastic
characters were previously thought to appear once in the history of
pterosaurs and may be correlated to the extension of the neck regions in
both groups. Because the homoplastic character states in the Azhdarchidae
and Ctenochasmatidae are limited to the mid-cervical vertebrae, these states
are termed convergent based on a definition of the term in a phylogenetic
context. A number of novel results from the analysis presented produce a
reorganization in the different species and taxa of the Pterodactyloidea.