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Coping w/ the rule; was metabolic rate [...]
Considering human design of complex systems, where
subsystems must work in concert to accomplish a given task; it is empirically
and theoretically unlikely that a human designer manifesting a novel system, or
even
refining an existing system, is going to be capable of building his
prototype(s)
at either end of the attainable size spectrum, even if such is his primary
intention. There
will inevitably be a period of testing and refinement, although forethought
and empirical knowledge can shorten it greatly. Conversely, lack of
forethought and
knowledge will likely increase time of optimization greatly, which leads to the
next point.
If the problems inherent to designing systems are a
fundamental aspect of reality, rather than mere subjective human experience;
and the process of evolution may be symbolized as a 'designer' w/out the
capability of forethought; then it follows that
the probability of a basal organism being well removed from the
minimum and maximum possible size for it's particular bodyplan/lifestyle is
quite high, even if there is no correlation of metabolic rate and evolutionary
rate to support Hazen's interesting and self-described "wild speculation". In
other words, basal organisms are likely to have 'room to move' in either
direction.
Further, here in the 'Newtonian zone', sometimes it is good to be small.
However, it can be effectively argued that whether engaged in competition for
resources, avoiding predation, or efficient utilization of resources obtained,
it is _usually_
advantageous to be bigger than the competition, within the limits inherent to
your particular design
and lifestyle.
Therefore, it doesn't seem remarkable that as basal organisms
radiate into more derived forms, many of them, probably a majority of
them, will increase in size. Given the vagaries of probability and game
theory, and the complexity of environment, it is also
not surprising that there are plenty of exceptions.
So my question is; isn't any "statistical appearance" of an evolutionary
undertow toward large size
that might be found in the fossil record easily explained by the foregoing
strictly practical considerations?
Don