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Panderichthys had tetrapod ears (today's Nature)
Nice stuff...
Martin D. Brazeau and Per E. Ahlberg. (2006) Tetrapod-like middle ear
architecture in a Devonian fish. Nature 439: 318-321.
Abstract: "Few fossils show the incipient stages of complex morphological
transformations. For example, the earliest stages in the remodelling of the
spiracular tract and suspensorium (jaw suspension) of osteolepiform fishes
into the middle ear of tetrapods have remained elusive. The most primitive
known tetrapods show a middle ear architecture that is very different from
osteolepiforms such as _Eusthenopteron_, with little indication of how this
transformation took place. Here we present an analysis of tetrapod middle
ear origins that is based on a detailed study of _Panderichthys_, the
immediate sister taxon of tetrapods. We show that the spiracular region is
radically transformed from osteolepiforms and represents the earliest stages
in the origin of the tetrapod middle ear architecture. The posterior
palatoquadrate of _Panderichthys_ is completely tetrapod-like and defines a
similarly tetrapod-like spiracular tract. The hyomandibula has lost its
distal portion, representing a previously unrecognized advance towards a
stapes-like morphology. This spiracular specialization suggests that the
middle ear of early tetrapods evolved initially as part of a spiracular
breathing apparatus."