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Re: New Titanosaur (Maxakalisaurus) from Brazil



Nope, Andesauridae has been recognized as a paraphyletic assemblage of basal titanosaurs. Of the "andesaurids" you mentioned, *Andesaurus* is the single non-titanosaurid titanosaur. In fact, one of the purported synapomorphies (presence of hyposphene-hypanthrum) has been shown to be lacking in *Argentinosaurus* (i.e., *Argentinosaurus* shares with other "titanosaurids"/lithostrotians, or derived titanosaurs, the character-state "hyposphene-hypanthrum absent)". *Epachthosaurus*, in turn, it is one of the most basal titanosaurid with procoelous mid and posterior caudal vertebrae (differing in this respect from *Andesaurus*, whose mid and posterior caudals are amphyplatyan). Indeed, it was chosen by Salgado (2003)
as a reference taxa in his restricted node-based definition of Titanosauridae (= Epachthosaurus sciuttoi + Saltasaurus loricatus).
Hope this helps,


Matías Soto

----- Original Message ----- From: "Jamie Stearns" <stearns5@cox.net>
To: <dinosaur@usc.edu>
Sent: Monday, August 28, 2006 9:28 PM
Subject: Re: New Titanosaur (Maxakalisaurus) from Brazil



Well, I see they took the time to classify this one somewhat, as I found titanosaur classification to be pretty difficult due to the number of described but still "incertae sedis" taxa out there. As far as I could tell, there were a number of basal titanosaurs, then Andesauridae which includes Epachthosaurus, Andesaurus, and Argentinosaurus (Is this still valid?), a proposed family of titanosaurs noted for (I think) amphicoelous vertebrae which includes Adamantisaurus, Rinconsaurus, Gondwanatitan, Laplatasaurus and possibly the original *Titanosaurus indicus*, the inclusion of which would make this the Titanosauridae, and finally the saltasaurids. I don't know exactly why the position of Aegyptosaurus is still uncertain, as it's been known for a long time now and Stromer most likely figured some of the (apparently somewhat complete) material in his papers somewhere. Another question: Is a skull known for Argentinosaurus? Many sources say there is no skull known, but I've also heard reports that a prospector named Daniel Eseisa had found the skull. The one on the Fernbank Museum's impressive restoration appears to be similar to a somewhat longer-faced Camarasaurus.

Moving on, I see another Diplodocus-like head has been added to this one. Was it preserved at all, or was it simply based on Rapetosaurus?


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