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Re: Tiktaalik
Titanosaurs across the board display opisthocoelous, procoelous,
amphicoelous and biconvex centra. The most parsimonious explanation
behind
this diversity in caudal morphology is that the front and back surfaces
could switch between one and the other conditions (e.g., amphicoelous or
opisthocoelous directly from procoelous), without always having to pass
through intermediate morphologies (e.g., weakly procoelous, amphiplatyan
conditions).
Hey, we're back to dinosaurs! I can't allow that... ;-)
In this case I don't have a better idea. However, gradual changes are also
suspected to occur. There are opisthocoelous salamanders in whose vertebrae
the ball contains a little socket in its center. Gradually enlarge that
socket, and you get a procoelous vertebra. (Without an amphicoelous or
amphiplatyan intermediate!) This is thought to be how the procoelous
salamanders (the Cretaceous Noterpetontidae... IIRC) came into being.