adult pterodactyloids (like _Quetzalcoatlus_) were flightless? What if they were fliers when young and segued into a more and more terrestrial niche as they grew? That would remove the problem of getting those gigantic things to take off.
I suppose a testable prediction would be adult forms more adapted to terrestriality than juvenile forms--I don't know if there's an appopriate ontogenetic series anywhere, though.