Ahlberg, P.E., Clack, J.A. and Blom, H. (2005). The axial skeleton of the Devonian tetrapod _Ichthyostega_. Nature 437: 137-140.
"[...] _Ichthyostega_ is the earliest vertebrate to show
obvious adaptations for non-swimming locomotion. Uniquely among early tetrapods, the presacral vertebral column shows pronounced regionalization of neural arch morphology, suggesting that it was adapted for dorsoventral rather than lateral flexion."
On another note...