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Fw: Applying Sereno's definitions to Neotetanurae Part 1



98 kB even though it's plain text! GAH! That's what I call work!

Martin, 1997

Tyrannoraptora
Maniraptoriformes
Eumaniraptora
|--Paraves
|  Maniraptora
|  Aves
|  |--Archaeopterygidae
|  |  Archaeopteryx
|  `--Ornithurae
|     Carinatae
|     Ichthyornis
`--Ornithomimiformes
  Oviraptoriformes
  |--all zero outgroup
  `--+--Dromaeosauridae
     |  |--Tyrannosauroidea
     |  |  Tyrannosaurus
     |  `--Deinonychus (+Dromaeosaurus?)
     `--+--Troodontidae
        |  Troodon
        `--+--Oviraptorosauria
           |  Oviraptor
           `--+--Alvarezsauridae
              |  Mononykinae
              |  Mononykus (+Shuvuuia)
              |--Ornithomimosauria
              |  Ornithomimus
              `--Struthiomimus

Ah, but no. Since he was too ignorant to correctly root his tree, we must do it for him:


+--allzero outgroup
`--Tyrannoraptora
  Maniraptoriformes
  Eumaniraptora
    |--Paraves
    |  Aves
    |    |--Archaeopterygidae
    |    |    *Archaeopteryx*
    |    `--Ornithurae
    |         Carinatae
    |         *Ichthyornis*
    `--Ornithomimiformes
       Oviraptoriformes
         |--Dromaeosauridae
         |    |--Tyrannosauroidea
         |    |  *Tyrannosaurus*
         |    `--*Deinonychus* ( + *Dromaeosaurus*?)
         `--+--Troodontidae
            |  *Troodon*
            `--+--Oviraptorosauria
               |  *Oviraptor*
               `--+--Alvarezsauridae
                  |  Mononykinae
                  |  *Mononykus*
                  |--Ornithomimosauria
                  |  *Ornithomimus*
                  `--*Struthiomimus*

It's Tyrannoraptora vs. Maniraptoriformes vs. Eumaniraptora again. This time we also have Maniraptora vs. Paraves, and Ornithomimiformes vs.
Oviraptoriformes. This again shows
Dromaeosauridae needs Tyrannosaurus as an external specifier.

Nothing of this changes.

Norell et al., 2001 and modifications.
This tree is taken from Makovicky et al. (2005), incorporating the new characters and three therizinosaurs from Kirkland et al. (2005) and the
modifications to Archaeopteryx and Rahonavis suggested by Mayr et al.
(2005).

Oho!

What modifications did they suggest to *Rahonavis*?

              `--+--Oviraptoriformes
                 |  |--Therizinosauria
                 |  |  Therizinosauroidea
                 |  |  |--Erlikosaurus (+Therizinosaurus?)
                 |  |  |--Segnosaurus
                 |  |  |--Alxasaurus
                 |  |  `--+--Nothronychus
                 |  |     |--Beipiaosaurus
                 |  |     `--Falcarius

Strange clade. Strikes me as... backwards... of course it's not a segnosaur analysis, but maybe it indicates that the segnosaurs belong somewhere else in the tree?


                       `--Dromaeosauridae
                          |--Buitreraptor
                          |--Unenlagiinae
                          |  |--Unenlagia
                          |  `--Rahonavis
                          |--Microraptorinae
                          |  |--Microraptor
                          |  `--Sinornithosaurus
                          `--+--Velociraptorinae
                             |  |--Velociraptor
                             |  |--IGM 100/1015
                             |  `--Deinonychus
                             `--Dromaeosaurinae
                                |--Saurornitholestes
                                |--Adasaurus
                                |--Achillobator
                                |--Utahraptor
                                `--Dromaeosaurus

So Dromaeosauridae is restored, and *Buitreraptor* and even *Rahonavis* stay inside...


Mayr et al., 2005

[...]
                 `--Ornithurae
                    |--Troodontidae

That hurts!

Chiappe and Walker, 2002
After deletion of many taxa, to provide structure to tree.

|--Archaeopteryx
`--+--Eoenantiornis
  |--Sinornis
  |--Eoalulavis
  |--Gobipteryx
  `--+--Neuquenornis
     |--Concornis
     `--Enantiornis

So not one of the names is applicable! :-D

Mortimer, in prep.
And here's my latest tree, a 90% consensus [...]

:-)

`--+--+--Fukuiraptor
  |  |--+--Labocania
  |  |  `--"Alashansaurus"
  |  `--+--Elmisaurus
  |     `--+--Megaraptor
  |        `--Tanycolagreus

A big-handed clade?

        |              `--+--Tarbosaurus
        |                 `--+--Bagaraatan
        |                    `--+--Aviatyrannis
        |                       `--Tyrannosaurus

Impressive :o)

                             `--Aves
                                |--Archaeopterygidae
                                |  |--+--Buitreraptor
                                |  |  `--Rahonavis
                                |  `--+--+--Jinfengopteryx
                                |     |  `--Scansoriopterygidae
                                |     `--+--Archaeopteryx
                                |        `--Wellnhoferia

Extremely well-behaved...

                                `--Ornithurae
                                   |--+--Sapeornis
                                   |  `--Omnivoropteryx
                                   `--+--Yandangornis
                                      `--+--Shenzhouraptor
                                         `--+--Confuciusornithidae
                                            |  |--Confuciusornis
                                            |  `--Changchengornis
                                            `--+--Jixiangornis
                                               `--+--Gobipteryx

Badly behaved!

`--Euornithes
  |--+--Eoalulavis
  |  `--Liaoningornis

That's normal.

`--+--Neuquenornis

That's normal for your analysis (I never got that, I think).

`--+--Iberomesornis

This is quite a performance! :-o

`--+--Boluochia

:-o
I guess its toothless pmx is among the reasons for this? I have this character in my matrix, but nevertheless *B.* stays in Enantiornithes (usually next to *Gobipteryx* for readily apparent reasons).


                       `--Carinatae
                          |--Ichthyornis
                          `--+--Hesperornithes
                             `--+--Iaceornis (+Passer?)
                                `--+--Hongshanornis
                                   `--"Archaeorhynchus"

These last two used to be the basalmost euorniths, and now they're among the most derived ones... :-o