[Date Prev][Date Next][Thread Prev][Thread Next][Date Index][Thread Index][Subject Index][Author Index]
RE: Heyuannia and Lu's monograph
--- dinosaur@usc.edu wrote:
> DINOSAUR Digest 3288
>
> Topics covered in this issue include:
>
> 1) Gorgosaurus sternbergi
> by Michael Lovejoy
> <michael@palaeoproductions.fsnet.co.uk>
> 2) Re: Gorgosaurus sternbergi
> by "Jaime A. Headden" <qilongia@yahoo.com>
> 3) Heyuannia and Lu's monograph
> by jay nair <sappororaptor@yahoo.com>
> 4) Re: Gorgosaurus sternbergi
> by Michael Lovejoy
> <michael@palaeoproductions.fsnet.co.uk>
> 5) RE: Heyuannia and Lu's monograph
> by Michael Mortimer <mickey_mortimer111@msn.com>
>
>
----------------------------------------------------------------------
>
> Date: Sat, 06 Aug 2005 12:55:41 +0100
> From: Michael Lovejoy
> <michael@palaeoproductions.fsnet.co.uk>
> To: DML <dinosaur@usc.edu>
> Subject: Gorgosaurus sternbergi
> Message-ID: <000501c59a7e$0532cfa0$6fc087d9@bob>
>
> Just a quick question: is AMNH 5664 considered
> Gorgosaurus sternbergi or an
> immature libratus?
> I noticed Currie used the species name sternbergi
> (very briefly!) in Cranial
> Anatomy of Tyrannosaurids from Alberta.
>
> So is it considered a valid, separate species?
>
> Regards,
> Michael Lovejoy
> Paleo Studio
>
>
> ------------------------------
>
> Date: Sat, 06 Aug 2005 07:07:10 -0700 (PDT)
> From: "Jaime A. Headden" <qilongia@yahoo.com>
> To: dinosaur@usc.edu
> Cc: michael@palaeoproductions.fsnet.co.uk
> Subject: Re: Gorgosaurus sternbergi
> Message-ID:
>
<20050806140711.97106.qmail@web33515.mail.mud.yahoo.com>
>
> Michael Lovejoy
> (michael@palaeoproductions.fsnet.co.uk) wrote:
>
> <Just a quick question: is AMNH 5664 considered
> Gorgosaurus sternbergi or an
> immature libratus? I noticed Currie used the species
> name sternbergi (very
> briefly!) in Cranial Anatomy of Tyrannosaurids from
> Alberta.>
>
> The taxon *G. sternbergi* is currently considered
> a junior synonym of
> *Gorgosaurus libratus* by Currie, and is listed in
> the noted paper in table 1
> as such a synonym.
>
> Cheers,
>
> Jaime A. Headden
>
> "Innocent, unbiased observation is a myth." --- P.B.
> Medawar (1969)
>
>
>
> ____________________________________________________
> Start your day with Yahoo! - make it your home page
> http://www.yahoo.com/r/hs
>
>
> ------------------------------
>
> Date: Sat, 06 Aug 2005 07:18:45 -0700 (PDT)
> From: jay nair <sappororaptor@yahoo.com>
> To: DML <dinosaur@usc.edu>
> Subject: Heyuannia and Lu's monograph
> Message-ID:
> <20050806141845.1914.qmail@web54404.mail.yahoo.com>
>
> Oviraptorid dinosaurs from Southern China
> Lu Junchang
>
> Geological Publishing House, Beijing.
> 2005. ISBN 7-116-04368-3. 200 pages + 8 plates.
> (In chinese: pp 1-83, including 36 figures & 3
> tables.
> In English: pp 85-200, including 5 geological
> figures.)
> All of the osteological figures are in the chinese
> section.
>
>
> Table of contents:
>
> Chinese section including bibliography
> pp 75-83 Bibliography
> English section
> pp 87 preface by Louis L. Jacobs
> 89 Abstract
> 90 Introduction
> 93 Geology of Heyuannia huangi
> 100 Osteology of Heyuannia huangi
> 100 Systematic Paleontology
> 112 Comparison and Discussion
> 116 Two new Oviraptorids from Nanxiong Basin
> 116 The Oviraptorid Material from Luyuan
> of Shixing County
> 119 An Oviraptorid Dinosaur from
> Xincheng Area of Nanxiong City
> 122 Phylogenetic Analysis of Oviraptorosauria
> 122 Materials and Methods
> 123 The Phylogenetic Analysis of
> Oviraptorosaurs Based on the
> Theropod Working group Matrix
> 130 The Phylogenetic Analysis of
> Oviraptorosaurs based on Modified
> Matrix
> 142 Conclusion
> 143 The Status of Oviraptorosauria Among
> Coelurosauria
> 144 Geographic and Stratigraphic Record of the
> Oviraptorosauria
> 147 Summary of Conclusion
> 149 Acknowledgements
> 151 Appendix A - for the first phylogenetic
> analysis. Character listes & character
> matices for each taxon
> 177 Appendix B - - for the second phylogenetic
> analysis. Character listes & character
> matices for each taxon
> 199 Plate Explanation
>
> Abstract
>
> Heyuannia huangi Lu, 2003 from the Late Cretaceous
> of
> Southern China is
> described in detail. The revised diagnosis of
> Heyuannia huangi includes the following characters:
> quadratojugal articular surface of the quadrate
> groove-lke, the quadrate diverticulum entering the
> quadrate
> anterolaterally, metacarpal I wrapped around
> metacarpal II proximally, pneumatic
> foramina present on the neural arches and ribs of
> cervical vertebrae, the angle between the scapula
> and
> coracoid approximately 145º, the ratio of
> coracoid length to scapular length approximately
> 0.35,
> the pubis as long as ischium, and the ratio of femur
> length to tibia length 0.8. Two new oviraptorids
> from
> the nanxiong Basin are also described. A
> phylogenetic
> analysis was
> carried out in two parts. The first phylogenetic
> analysis is to test the systematic position of
> Oviraptorosauria among ceolurosaurid dinosaurs. This
> analysis shows that Oviraptorosauria form a
> monophyletic group. Incisivosaurus
> gauthieri is the most basal oviraptorosaur and
> Caudipteryx is the second most primitive form.
> Alvarezsauridae is a sister group to TO
> (Therizinosauroidea and oviraptorosauria). Within
> this
> more inclusive analysis (among Ceolurosauria), all
> the
> derived forms of Oviraptorosauria (excluding the
> Primitive forms such as Incisivosaurus gauthieri and
> Caudipteryx zoui) form an unresolved polytomy. The
> second analysis is to determine the
> interrelation-ships among oviraptorosaurs. This less
> inclusive analysis (only a few groups of
> Ceolurosauria
> were employed for analysis) allows better resolution
> within Oviraptorosauria. The analysis shows
> that all oviraptorosaurs form a monophyletic group,
> Incisivosaurus gauthieri is the most basal
> oviraptotosaur, Caudipteryx zoui is the second most
> primitive form and Avimimus portentosus is basal to
> the derived oviraptorosaurs. The two new taxa from
> Nanxiong Basin are not nested within same clade.
> Specimen BPV-112 may be the first caenagnathid
> oviraptorosaur found from
> southern China.
>
> Notes
>
> Lu describes the osteology of Heyuannia in detail.
> The description is based on the type specimen (the
> most complete specimen) and at least 3 other
> individuals. The incomplete skull was most likely
> crestless (like Ingenia, Conchoraptor & Khaan). The
> lower jaw is similar to other oviraptorids (being
> short & deep). Heyuannia has more cervicals &
> sacrals
> than most other small non-avain theropods. There are
> 13 cervicals in Heyuannia (also Richenia). Ingenia
> has
> 11 while Caudipteryx has 10. There are 8 sacrals
> (Negmetia & Ingenia have 8, Nominga 5 and Richenia
> 7).
>
> There are 3 sternal ribs in Heyuannia. This may be a
> derived character or a result of non-preservation in
> other theropods (Dromaeosauridae & Aves have 3+).
>
> The manus is at 60 degrees to the forearm. The first
> digit is very strong in Heyuannia - similar to
> Ingenia. The dorsal margin of the ilium is straight
> (as in Ingenia & Nemegtia). The tibia/femur ratio is
> similar to many small theropods & flightless birds.
> The distal end of the fibula reaches the tarsals,
> unlike in Ingenia. The ascending process (of the
> astragulus) is thin & tapers up.
>
> Lu mentions two other oviraptorosaurs.
>
> The first (BPV-112) is based on a nearly complete
> sacrum, 3 posterior dorsals, 3 articulated anterior
> caudals, 2 ilia and portions of the hindlimbs.
> Lu refers it to Caenagnathidae gen et sp indet.
> BPV-112 is a new taxon, but isn't named because of
> the
> scarce materials. Phylogenetically, it sits closest
> to
> Chirostenotes in Lu's analyses.
> Two autopomorphies include the higher ratio of the
> height/length of the ilium (not so convincing
> considering the variability of ratios in specimens
> of
> Ingenia) and the presense of a large foramen on the
> proximal and of the femur.
> I think BPV-112 maybe is the first definite
> caenagnathid from China. It's from the Maastrichtian
> Pingling Formation.
>
> The second unnamed oviraptorid is unnumbered. It is
> specimen E-1 at the Nanxiang Musuem, so Lu
> temporarily
> calls it NXMV.
> NXMV is the rear half of a skull & nearly complete
> lower jaw, 2 cervicals, 9 dorsals & an incomplete
> right pes. Originally thought to be from Paleocene
> deposits, a Late Cret age is shown from K-Ar dating
> (Pingling Member of the Shanghu Formation).
> So NXMV may prove to be the youngest known member of
> the Oviraptoridae. Lu also decides not to name it
> because it's still in preparation. Expect a more
> detailed publication with a name for NXMV from Lu in
> the future.
> The lower jaw seems closer to Oviraptor
> (philoceratops) in its proportions. The vertebrae
> have
> well developed pleurocoels. NXMV is nested closest
> to
> Heyuannia & Nemegtia in Lu's analysis.
>
>
> Lu conducted 2 analyses. The first to test the
> position of Oviraptorosauria among other theropods
> including birds (using Hwang et al's ''Theropod
> working group'' matrix). The second was to determine
> the interelationships of oviraptorosaurs (using
> Maryanska et al's matrix)
> Characters used to diagnose Caudipteryx dongi seem
> to
> be related to ontogeny, so Lu sinks C. dongi into C.
> zoui.
>
> The most interesting outcome of Lu's strict
> concensus
> tree is that Alvarezsauridae form the sister taxon
> to
> [Therizinosauroidea + Oviraptorosauria] - far away
> from a relatively 'basal' Ornithomimosauria.
> >From fig 18:
>
> Allosaurus (Sinraptor + (Tyrannosauroidea +
> (Ornithomimosauria + (Ornitholestes +
> (Compsognathidae
> + ([Aves + Deinonychusauria] + [Alvarezsauridae +
> Oviraptorosauria/Therizinosauria]
>
> The second analysis found the following
> interelationships
>
> 1)= a monophyletic Oviraptorosauria (Eighteen
> synapomorphies). Includes Incisivosaurus and other
> (2)
> taxa more derived than Incisivosaurus
>
> 2)= Caudipteryx and other (3) taxa more derived than
> Caudipteryx. (eight synapomorphies)
>
> 3)= Microvenator + (4). Five synapomorphies
>
> 4)= Avimimus + (5). Three synapomorphies
>
> 5)= Caenagnathoidea. (Oviraptoroidea of Barsbold
> 1976.
> Caenagnathoidea (Sternberg, 1940) has priority.
> Caenagnathoidea is Oviraptoridae (6) (Barsbold,
> 1976)
> + Caenagnathidae (Sternberg, 1940)
>
> Caenagnathidae includes BPV-112 and Chirostenotes,
> and
> possibly Caenagnathasia
>
> 6)=Oviraptoridae includes (7)[Ingeniinae,
> Oviraptorinae] & Khaan
>
> 7)= Ingeniinae + Oviraptorinae, excludes Khaan
>
> Ingeniinae includes Heyuannia, Nemegtia, Citipati,
> Richenia, Nomingia & NXMV
> Within Ingeniinae, Nomingia + Richenia form a
> derived
> group with Heyuannia, Nemegtia, NXMV, Citipati,
> Ingenia forming successive outgroups to the derived
> [Nomingia + Richenia] in ascending order.
>
> Oviraptorinae includes Conchoraptor, Oviraptor & GIN
> 10042 (GIN 10042 has been previously discussed on
> the
> DML)
> Within Oviraptorinae, Lu weakly finds GIN 10042
> closest to Oviraptor, with Conchoraptor the
> outgroup.
>
>
> Sappororaptor
>
>
> Refs
>
> Barsbold. 1976 On a new Late Cretaceous family of
> small theropods (Oviraptoridae fam. n.) of Mongolia.
> (in Russian). Doklady Akademii Nauk SSSR 226:
> 685-688.
>
> Hwang, Norell, Ji, and Gao. 2004. A large
> compsognathid from the Early Cretaceous Yixian
> Formation of China. Jounal of Systematic
> Palaeontology
> 2(1): 13-30.
>
> Maryanska, Osmolska, and Wolsan. 2002. Avialan
> status
> of Oviraptorosauria. Acta Pal. Pol 47: 97-116
>
> Sternberg. 1940. A toothless bird from the
> Cretaceous
> of Alberta. Journal of Pal. 14: 81-85
>
>
>
>
>
>
> __________________________________________________
> Do You Yahoo!?
> Tired of spam? Yahoo! Mail has the best spam
> protection around
> http://mail.yahoo.com
>
> ------------------------------
>
> Date: Sat, 06 Aug 2005 19:22:04 +0100
> From: Michael Lovejoy
> <michael@palaeoproductions.fsnet.co.uk>
> To: qilongia@yahoo.com, dinosaur@usc.edu
> Subject: Re: Gorgosaurus sternbergi
> Message-ID: <006901c59ab3$c0afd720$a5e14c51@bob>
>
> Thanks very much for pointing that out! I'm really
> going to have to learn to
> read properly...
> Regards,
> Michael Lovejoy
> Paleo Studio
>
> ----- Original Message -----
> From: "Jaime A. Headden" <qilongia@yahoo.com>
> To: <dinosaur@usc.edu>
> Cc: <michael@palaeoproductions.fsnet.co.uk>
> Sent: Saturday, August 06, 2005 3:07 PM
> Subject: Re: Gorgosaurus sternbergi
>
>
> > Michael Lovejoy
> (michael@palaeoproductions.fsnet.co.uk) wrote:
> >
> > <Just a quick question: is AMNH 5664 considered
> Gorgosaurus sternbergi or
> an
> > immature libratus? I noticed Currie used the
> species name sternbergi (very
> > briefly!) in Cranial Anatomy of Tyrannosaurids
> from Alberta.>
> >
> > The taxon *G. sternbergi* is currently
> considered a junior synonym of
> > *Gorgosaurus libratus* by Currie, and is listed in
> the noted paper in
> table 1
> > as such a synonym.
> >
> > Cheers,
> >
> > Jaime A. Headden
> >
> > "Innocent, unbiased observation is a myth." ---
> P.B. Medawar (1969)
> >
> >
> >
> >
> ____________________________________________________
> > Start your day with Yahoo! - make it your home
> page
> > http://www.yahoo.com/r/hs
> >
> >
> >
> >
> > --
> > No virus found in this incoming message.
> > Checked by AVG Anti-Virus.
> > Version: 7.0.338 / Virus Database: 267.10.0/63 -
> Release Date: 03/08/2005
> >
> >
>
>
> ------------------------------
>
> Date: Sat, 06 Aug 2005 19:39:25 +0000
> From: Michael Mortimer <mickey_mortimer111@msn.com>
> To: dinosaur@usc.edu
> Subject: RE: Heyuannia and Lu's monograph
> Message-ID:
> <BAY106-F15DD66784C4319C5E0EB61ACC60@phx.gbl>
>
> jay nair wrote-
>
> >Lu mentions two other oviraptorosaurs.
> >
> >The first (BPV-112) is based on a nearly complete
> >sacrum, 3 posterior dorsals, 3 articulated anterior
> >caudals, 2 ilia and portions of the hindlimbs.
> >Lu refers it to Caenagnathidae gen et sp indet.
> >BPV-112 is a new taxon, but isn't named because of
> the
> >scarce materials. Phylogenetically, it sits closest
> to
> >Chirostenotes in Lu's analyses.
> >Two autopomorphies include the higher ratio of the
> >height/length of the ilium (not so convincing
> >considering the variability of ratios in specimens
> of
> >Ingenia) and the presense of a large foramen on the
> >proximal and of the femur.
> >I think BPV-112 maybe is the first definite
> >caenagnathid from China. It's from the
> Maastrichtian
> >Pingling Formation.
>
> Funny how people don't understand the concept of a
> specimen being
> indeterminate. If it's a new taxon with
> autapomorphies, it can't be gen. et
> sp. indet.. Lu actually names this in his thesis
> (which I'm betting is
> almost identical to this book). When I ran my
> analysis back in May, it came
> out as the sister group of Caenagnathoidea.
No, he doesn't name it in the published volume.
But you're right about the book being the same as his
thesis - nearly. He states this in his
acknowledgements.
> >The most interesting outcome of Lu's strict
> concensus
> >tree is that Alvarezsauridae form the sister taxon
> to
> >[Therizinosauroidea + Oviraptorosauria] - far away
> >from a relatively 'basal' Ornithomimosauria.
>
> This happened in the first Theropod Working Group
> published analysis too.
> Back in the Ostrom volume.
>
> Graphically, Lu's phylogeny is-
>
> |--Incisivosaurus
> `--+--Caudipteryx
> `--+--Microvenator
> `--+--Avimimus
> `--+--+--BPV-112
> | `--Chirostenotes
> `--+--Khaan
> `--+--+--Conchoraptor
> | `--+--Oviraptor
> | `--GIN 100-42
> `--+--Heyuannia
> `--+--Nemegtia
> `--+--NXMV
> `--+--Citipati
> `--+--Ingenia
> `--+--Rinchenia
> `--Nomingia
>
> Note he had to exclude Herrerasaurus, Coelophysis
> and Confuciusornis from
> Maryanska et al.'s matrix to achieve this result.
> When the former two are
> included, Avimimus is the most basal, Microvenator
> is sister to
> Chirostenotes+BMV-112, and Oviraptor is the
> basalmost caenagnathoid.
>
> Compare this to my results as of May-
> |--Incisivosaurus
> `--+--Caudipteryx
> `--+--Avimimus
> `--+--Nomingia
> `--+--BPV-112
> `--+--Caenagnathidae
> `--+--Microvenator
> `--+--Oviraptor
> `--+--Conchoraptor
> `--+--Khaan
> `--+--+--Nemegtia
> | `--Heyuannia
> `--+--Citipati
> (includes GIN 100-42)
> `--+--+--ZPAL
> MgD-I/95
> | `--Ingenia
> `--+--Rinchenia
> `--NXMV
>
> There are definitely some similarities. They both
> share the same basic
> topology-
> |--Incisivosaurus
> `--+--Caudipteryx
> `--+--Avimimus
> `--+--BPV-112
> |--Caenagnathidae
> `--+--Oviraptor
> |--Conchoraptor
> |--Khaan
> `--+--Heyuannia
> |--Nemegtia
> `--+--Citipati
> `--+--Ingenia
> `--Rinchenia
>
> Since May, I've added many codings for Nemegtia, but
> I'm in the process of
> reformatting the drive on the computer with PAUP, so
> I can't see the
> codings' effect yet.
>
> Mickey Mortimer
>
>
>
____________________________________________________
Start your day with Yahoo! - make it your home page
http://www.yahoo.com/r/hs