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Re: Benton and Kinman (long)




Thanks David,
Benton's 1997 classification is a valiant effort to store cladistic information, while at the same time recognizing some paraphyletic groups. He obviously recognizes Crocodylia sensu lato and separates Aves from Reptilia. But as much as admire his effort, it is still too messy (and listing a Superclass among a list of Orders is a little confusing---although I clearly understand why he did so):


Infraclass Rhipidistia
 Order Porolepiformes
 Order Rhizodontiformes
 Order Osteolepiformes
 Order Panderichthyida
 Superclass Tetrapoda

In my 1994 classification, I recognized the same clade informally through a code sequence within Class Sarcopterygea (an abbreviated version as follows, showing how my markers differ from his):

5  Sarcopterygea
    1  .....
         etc.
    4  Porolepiformes
    5  Osteolepiformes
    6  Rhizodontiformes
    7  Panderichthyiformes
    8  {{Amphibea}} (thence to Reptilea,
         Avea and Mammalea) (= Tetrapoda)

_1_ Amphibea
     1   Ichthyostegiformes
         ....etc.
     2D  Diadectiformes
     2E  {{Reptilea}} (thence to Avea
           and Mammalea) (= Amniota)
     3   Aistopodiformes
         ....... etc.

Note that I have dropped "rank" names by completing the trend toward standardized endings above family level---but this is very controversial, and the system works using the names with traditional endings (Mammalia, Aves, etc.), so don't let that put you off too much (I still use Mammalia and Aves in my day-to-day correspondence, but standardized endings may catch on in the future).
By the way, the clade Sarcopterygea 5-8 is what some cladists call Tetrapodomorpha, and this same coded clade will remain even if Rhizodontiformes split off before Osteolepiformes (I will just have to switch the order of those two Orders if that topology is confirmed).
Anyway, Benton is trying to accomplish the same thing I am, but using (IMHO) old-fashioned intermediate ranks in a manner somewhat similar to the strict cladist McKenna (in mammalian classifications). Benton is a great improvement over McKenna, but still a little too messy for my tastes. If you don't like my approach, I very, very highly recommend the way Benton does it. Whether Benton chooses to adopt any of my conventions in future classications remains to be seen, but we obviously share very similar Ashlockian (centrist) tendencies.
------Ken Kinman


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