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Re: seeking clarification on the cladism debate (RE: hidden "cladistic" ranks)




If I understand David's question correctly (about "setting points" along a phylogeny), this is precisely what I do in my classifications. I do try to minimize the use of paraphyletic groups, cutting the Tree of Life at only certain strategic "points" which are very biological significant (and which often have long histories in the biological literature, thus promoting some degree of continuity and relative stability when possible).
At such a "re-setting" point, a divergent and rapidly evolving taxon is bumped up a rank (sometimes even more than one rank). In my dinosaur classification, Family Dromaeosauridae is presently coded as sister group to Class Aves (the latter having thus been "promoted" by two ranks). This two-rank jump is shown by two separate {{Aves}} markers: (1) one at family level (coded as sister group to Dromaeosauridae) within the classification of Order Saurischiformes; and (2) one at order level (within Class Reptilia) just after Saurischiformes, coded to show that Class Aves has been paraphyletically removed from Order Saurischiformes (and thus has a sister group within it), and therefore from Class Reptilia as well.
This kind of cross-referencing system thus renders paraphyletic groups informationally "complete", by adding a marker for the "exgroup" which has been promoted to higher rank. Since "markered" taxa are no longer incomplete in informationally terms, I call them semi-paraphyletic (or sometimes call them "semi-holophyletic" to try and make it more palatable to strict cladists).
Thus unlike traditional "eclectic" classifications, paraphyletic groups are clearly marked as such, and sister group information is retained. And it does this without the adverse "side-effects" of purely cladistic classifications (e.g., multiplicity of names, hierarchical instability, lack of anagenetic information, and decreases in utility). In many ways, we can have our cake and eat it too (i.e. get the best of both cladistic and traditional approaches), and avoid many of the drawbacks of both. The Kinman System is therefore more synergistic than a mere compromise (and it certainly is not like being half pregnant, which is something I hear all too often).
I therefore completely disagree with Mike's statement that nesting a higher ranked taxon in one of lower rank renders such ranks meaningless. On the contrary, it is a more efficient and stable way of reflecting the punctuated-equilibrium irregularities of evolutionary history, particularly those which are magnified by mass extinctions. No classification can truly be devoid of subjectivity, but we can certainly try to minimize such subjectivity. Instead of eliminating subjectivity, "strictly" cladistic classifications unfortunately just substitute new forms of subjectivity for the old (and it comes at a very high price-----as the side-effects mentioned above attest).
----------Ken Kinman
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David Elliott wrote:
> Has the idea of setting points along phylogenies where the ranks are "reset" been discussed?

Mike Keesey responded:
That would undermine the entire concept of an absolute rank. A taxon with a lower level rank (e.g. Ordo Saurischia) should never include a taxon with a higher level rank (e.g. Classis Aves). If it can, then the ranks are rendered even more meaningless than before.

Furthermore, picking these points would be an entirely subjective practice, without any scientific principle to back it up.


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