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Re: hypocleidium (an interclavicle???)



WOW,
So you are saying that the theropod "hypocleidium" and the pygostylian hypocleidium may not be homologous (and therefore should probably be coded as separate characters)? If so, is there any way to know which of these types is present in Protoavis??
I didn't know that the "hypocleidium" (the theropod type) was present in allosauroids or tyrannosauroids. Aren't such structures uncommon among non-avian theropods, or perhaps they are common but just not often in ossified form?
Thanks for the info, Ken
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Jaime A. Headden wrote:
No memeber of the Dinosauromorpha possesses an interclavicle. This is a distinct osseous element
that interjects between coracoids and between each of the clavicle bones; the interclavicle has a
tri-radiate morphology, essentially Y-shaped in form, leading to the confusion I'm sure that spawned the question. In the period of time that the dinosaurian furcula of *Oviraptor* was offered as an interclavicle since Osborn's 1924 description, it has since been the prevalent
scientific conclusion that this element is in fact a fusion or collection of two clavicles without
an interclavicle, and no the interclavbicle itself. Even as recent as 1986, the fusion of the clavicles was regarded as "fused clavicles" and not a "furcula," and is still regarded as such by
the more resistant of the ornithologists (see Feduccia, 1996, for example).


The hypocleidium is a structure that appears early in the development of theropods with furculae, appearing in allosauroids and tyrannosauroids, as a distinct process. It appears to be an extension of the fusion of two diaphyses, such as the ramal shafts of the paired clavicles. It
is lost in early birds, such as *Archaeopteryx* and *Confuciusornis*, but is derived in higher
groups, so that nearly all modern birds possess it. Rather than re-expression, it is likely that
the hypocleidium in higher birds is probably a neomorph structure adapted based on tendon
attachment, rather than the ends of two fused diaphyses.



===== Jaime A. Headden

  Aaaaaaaaaaaaaaaaaaaahhhhhhhhhhhhhhhhhhr-gen-ti-na
  Where the Wind Comes Sweeping Down the Pampas!!!!

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