Re: Pterosaur Help

[Dinogeorge@aol.com]

In a message dated 11/23/01 9:10:22 PM EST, mhanson54@home.com writes:

<< Recently, I have been trying to compile a list of pterosaur species. I
 really need help with Ornithocheirus! It is said that there have been 36
 species of Ornithocheirus named, and so far, I have only found 15 and found
 almost none of their describers and years of description. I would greatly
 appreciate it if someone could help me with finding all attributed species,
 describers, years of descriptions, and synonymized genera. I would also
 appreciate some help with the following genera; Nyctosaurus, Pterodactylus,
 Rhamphorhynchus, Germanodactylus, Scaphognathus, Cycnorhamphus, Dimorphodon,
 and Campylognathoides. I have noticed that the more well-known pterosaurs
 are the ones I have difficulty with compiling complete lists.
     Thanks,
           Michael >>

OK, here's a listing for Ornithocheiridae. It's a little out of date, but 
help yourself. There are more species of Ornithocheirus scattered throughout 
Pterosauria, but this should get you started:

Family: ORNITHOCHEIRIDAE (nom.    emend. ex Ornithocheirae) Seeley, 1870
Census: 6 genera, 34 species (19 doubtful)

    = Ornithocheirinae Williston, 1897
    = Ornithocheiroidea Williston, 1897
    = Ornithocheiridae Zittel & Eastman, 1902
    = Ornithocheridae Nopcsa, 1923 [sic]

Genus: Araripesaurus Price, 1971
A. castilhoi Price, 1971â

Genus: Arthurdactylus Frey & Martill, 1994
A. conandoylei Frey & Martill, 1994 emend nov.â
= Arthurdactylus conan-doylei Frey & Martill, 1994*

Genus: Brasileodactylus Kellner, 1984
B. araripensis Kellner, 1984â
= Brasileodactylus araripendis Kellner, 1989 [sic]

Genus: Lonchodectes Hooley, 1914
?L. giganteus (Bowerbank, 1846) [nomen dubium]
= Pterodactylus giganteus Bowerbank, 1846 non (Oken, 1819) [nomen dubium]
= Ornithocheirus giganteus (Bowerbank, 1846) Newton, 1888 [nomen dubium]
= Ornithodesmus giganteus (Bowerbank, 1846) Kuhn, 1967 [nomen dubium]
= Pterodactylus conirostris Owen vide Dixon, 1850 [nomen dubium]
= Ornithochirus giganteus Lydekker, 1888 [sic]
L. compressirostris (Owen, 1851) Hooley, 1914
= Pterodactylus compressirostris Owen, 1851
= Ornithocheirus compressirostris (Owen, 1851) Seeley, 1870
= Ornithochirus compressirostris Lydekker, 1888 [sic]
= Ornithocheirus compressiformis Gaal, 1926 [sic]
?L. machaerorhynchus (Seeley, 1864) [nomen dubium]
= Pterodactylus machaerorhynchus Seeley, 1864 [nomen dubium]
= Ornithocheirus machaerorhynchus (Seeley, 1864) Seeley, 1870 [nomen dubium]
= Ptenodactylus machaerorhynchus (Seeley, 1864) Seeley, 1869 [nomen dubium]
= Pterodactylus machaerhynchus Sauvage, 1882 [sic]
= Ornithocheirus machaeorhynchus Hooley, 1914 [sic]
?L. oweni (Seeley, 1864) [nomen dubium]
= Pterodactylus oweni Seeley, 1864 [nomen dubium]
= Ornithocheirus oweni (Seeley, 1864) Seeley, 1870 [nomen dubium]
= Ptenodactylus oweni (Seeley, 1864) Seeley, 1869 [nomen dubium]
= Pterodactylus owenii Sauvage, 1882 [sic]
?L. microdon (Seeley, 1870) [nomen dubium]
= Ornithocheirus microdon Seeley, 1870 [nomen dubium]
= Pterodactylus microdon (Seeley, 1870) Sauvage, 1882 [nomen dubium]
= Ptenodactylus microdon Seeley, 1869 [nomen nudum]
?L. scaphorhynchus (Seeley, 1870) [nomen dubium]
= Ornithocheirus scaphorhynchus Seeley, 1870 [nomen dubium]
= Pterodactylus scaphorhynchus (Seeley, 1870) Sauvage, 1882 [nomen dubium]
= Ptenodactylus scaphorhynchus Seeley, 1869 [nomen nudum]
?L. tenuirostris (Seeley, 1870) [nomen dubium]
= Ornithocheirus tenuirostris Seeley, 1870 [nomen dubium]
= Pterodactylus tenuirostris (Seeley, 1870) Sauvage, 1882 [nomen dubium]
= Ptenodactylus tenuirostris Seeley, 1869 [nomen nudum]
?L. daviesii (Owen, 1874)
= Pterodactylus daviesii Owen, 1874
= Ornithocheirus daviesii Owen, 1874 emend. Olshevsky, 1991
= Ornithocheirus daviesi (Owen, 1874) Newton, 1888*
= Pterodactylus daviesii Owen, 1874 emend. Olshevsky, 1991
= Pterodactylus daviesi (Owen, 1874) Sauvage, 1882*
= Ornithochirus daviesi Lydekker, 1888 [sic]
?L. sagittirostris (Owen, 1874) Unwin vide Benton, 1993
= Pterodactylus sagittirostris Owen, 1874
= Ornithocheirus sagittirostris (Owen, 1874) Newton, 1888
= Ornithodesmus sagittirostris (Owen, 1874)
NOTE: Generally considered a synonym of Ornithocheirus, this genus is 
referred to an unnamed new family by Unwin (in Benton, ed., 1993). It is here 
referred to Ornithocheiridae provisionally. See also note for Ornithocheirus.

Genus: Ornithocheirus Seeley, 1869
= Cimoliornis Owen, 1846
= Cretornis Fritsch, 1880 [nomen dubium]
= Lithosteornis Gervais, 1844 [nomen nudum]
= Ornithochirus Lydekker, 1888/Cope, 1872 [sic]
= Osteornis Gervais, 1844 [nomen nudum]
= Palaeornis Mantell, 1835/Vigors, 1825
= Ptenodactylus Seeley, 1869 (preoccupied?)
?O. clifti (Mantell, 1835) Newton, 1888
= Palaeornis clifti Mantell, 1835â
= Pterodactylus clifti (Mantell, 1835) Bronn, 1848
= Osteornis ardeaceus Gervais, 1844 [nomen nudum]â
= Lithosteornis ardeaceus (Gervais, 1844) [nomen nudum]â
= Pterodactylus silvestris Owen, 1845
= Pterodactylus sylvestris Owen, 1859 [sic]
= Ornithochirus clifti Lydekker, 1888 [sic]
?O. diomedeus (Owen, 1846) Newton, 1888
= Osteornis diomedeus Owen, 1844 [nomen nudum]
= Cimoliornis diomedeus Owen, 1846â
= Pterodactylus diomedeus (Owen, 1846) Owen, 1851
= Pterodactylus diomedius (Owen, 1846)
= Pterodactylus giganteus Morris, 1854 non (Oken, 1819) non Bowerbank, 1846
= Cimoliornis diomedius Lydekker, 1888 [sic]
= Ornithochirus diomedius Lydekker, 1888 [sic]
= Ornithocheirus diomedius Hooley, 1914 [sic]
O. fittoni (Owen, 1858) Seeley, 1870
= Pterodactylus fittoni Owen, 1858
= Ptenodactylus fittoni (Owen, 1858) Seeley, 1869
= Ornithochirus fittoni Lydekker, 1888 [sic]
O. sedgwickii (Owen, 1859) emend. Newton, 1888
= Ornithocheirus sedgwicki (Owen, 1859) Seeley, 1870*
= Pterodactylus sedgwickii Owen, 1859 (not 1858)
= Coloborhynchus sedgwickii (Owen, 1859) Owen, 1874
= Ptenodactylus sedgwickii (Owen, 1859) emend. nov.â
= Ptenodactylus sedgwicki (Owen, 1859) Seeley, 1869â*
= Coloborhynchus sedgwicki Sauvage, 1882 [sic]
= Pterodactylus sedgwicki Sauvage, 1882 [sic]
= Ornithochirus sedgwicki Lydekker, 1888 [sic]
?O. macrorhinus (Seeley, 1869) [nomen nudum]
= Ptenodactylus macrorhinus Seeley, 1869 [nomen nudum]
= Pterodactylus macrohinus Sauvage, 1882 [sic]
O. brachyrhinus Seeley, 1870 [nomen dubium]
= Pterodactylus brachyrhinus (Seeley, 1870) Sauvage, 1882 [nomen dubium]
= Ptenodactylus brachyrhinus Seeley, 1869 [nomen nudum]
O. colorhinus Seeley, 1870 [nomen dubium]
= Pterodactylus colorhinus (Seeley, 1870) Sauvage, 1882 [nomen dubium]
= Ptenodactylus colorhinus Seeley, 1869 [nomen nudum]
?O. curtus (Owen, 1870) Newton, 1888 [nomen dubium]
= Pterodactylus curtus Owen, 1870 [nomen dubium]
= Ornithochirus curtus Lydekker, 1888 [sic]
O. dentatus Seeley, 1870 [nomen dubium]
= Pterodactylus dentatus (Seeley, 1870) Sauvage, 1882 [nomen dubium]
= Ptenodactylus dentatus Seeley, 1869 [nomen nudum]
O. denticulatus Seeley, 1870 [nomen dubium]
= Ornithochirus denticulatus Lydekker, 1888 [sic]
O. enchorhynchus Seeley, 1870 [nomen dubium]
= Pterodactylus enchorhynchus (Seeley, 1870) Sauvage, 1882 [nomen dubium]
= Ptenodactylus enchorhynchus Seeley, 1869 [nomen nudum]
O. huxleyi Seeley, 1870 [nomen dubium]
O. nasutus Seeley, 1870 [nomen dubium]
= Pterodactylus nasutus (Seeley, 1870) Sauvage, 1882 [nomen dubium]
= Ptenodactylus nasutus Seeley, 1869 [nomen nudum]
O. oxyrhinus Seeley, 1870 [nomen dubium]
= Pterodactylus oxyrhinus (Seeley, 1870) Sauvage, 1882 [nomen dubium]
= Ptenodactylus oxyrhinus Seeley, 1869 [nomen nudum]
O. polyodon Seeley, 1870 [nomen dubium]
= Ptenodactylus polyodon Seeley, 1869 [nomen nudum]
= Pterodactylus platyodon Sauvage, 1882 [sic]
O. xyphorhynchus Seeley, 1870 [nomen dubium]
= Ornithochirus xyphorhynchus (Seeley, 1870) [nomen dubium]
?O. hlavaci (Fritsch, 1880) Fritsch, 1905 [nomen dubium]
= Ornithocheirus hlavà i (Fritsch, 1880) Fritsch, 1905 [nomen dubium]*
= Cretornis hlavaci Fritsch, 1880 [nomen dubium]â
= Cretornis hlavà i Fritsch, 1880 [nomen dubium]â*
= Cretornis hlavatschi Lydekker, 1888 [sic]
= Ornithocheirus hlavatschi Newton, 1888 [sic]
= Ornithochirus hlavatschi Lydekker, 1888 [sic]
?O. bunzeli Seeley, 1881 emend. Sauvage, 1882
= Ornithocheirus bÃnzeli Seeley, 1881*
= Ornithocheirus bunzelii Sauvage, 1882 [sic]
NOTE: This species is noted as a possible azhdarchid by Unwin (pers. comm. in 
Nessov, 1991).
?O. hilsensis Koken, 1883 [nomen dubium]
NOTE: This species probably represents a small theropod dinosaur (Meyer, 
1884; Williston, 1885).
O. wiedenrothi Wild, 1990
NOTE: This genus has become a terrible wastebasket for miscellaneous 
pterosaurian remains from the Cambridge Greensand and elsewhere. Seeley 
failed to nominate a type species until 1881, when he noted it was 
Ornithocheirus simusâthe first species listed under the genus by Seeley in 
1869. But by then Owen had already made Pterodactylus simus the type species 
of Criorhynchus (Owen, 1874). The "type" species often cited by 20th-century 
authors (e.g., Kuhn, 1967; Wellnhofer, 1978; Wild, 1990), Ornithocheirus 
compressirostris, was definitely separated from Ornithocheirus into a 
different genus, Lonchodectes, by Hooley (1914). Unfortunately, in his review 
of Seeley's numerous species, Hooley himself failed to specify type species 
for his genera Amblydectes and Lonchodectes; nor did he establish a neotype 
species for Ornithocheirus. Pending publication of revisory work by D. Unwin, 
the various species of Ornithocheirus are organized here according to Hooley 
(1914) and those parts of Unwin's reclassification that have already been 
published (as in the section on Pterosauria in The Fossil Record 2 [Benton, 
ed., 1993]). No type species is indicated here for this genus.
The genus Ptenodactylus, created by Seeley in 1869 for numerous remains in 
the Woodwardian Museum, lacked a type species until the aforementioned 1881 
work, when he noted that its numerous species were "allied with Pterodactylus 
sedgwicki," thus making that speciesâthe first listed by him in 1869âthe 
type of the genus. But already in 1870 Seeley had abandoned Ptenodactylus (as 
preoccupied?: Lydekker, 1888) in favor of Ornithocheirus as the generic name 
for all the Woodwardian Museum species.

Genus: Santanadactylus de BuisonjÃ, 1980
= Sandactylus Bowler, 1989 [sic]
S. brasilensis de BuisonjÃ, 1980â
= Santandactylus brasiliensis Nessov, 1991 [sic]
?S. araripensis Wellnhofer, 1985
?S. pricei Wellhofer, 1985
?S. spixi Wellhofer, 1985
NOTE: Bennett (1989) considered Santanadactylus araripensis and S. pricei to 
be pteranodontids and S. spixi to be a dsungaripterid. Bennett also noted 
that the type species may be based on a composite specimen: part 
pteranodontid, part something else. Wellnhofer (1991) retains the former two 
species in this genus but regards S. spixi as Pterodactyloidia incertae 
sedis. Kellner & Campos (1992) regard Santanadactylus spixi as a tapejarid.