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Nicholas Gardner wrote-
Below is my rather early and tenative classification of Maniraptora
(Passer + Enigmosaurus) that includes
tyrannosaurs.
Maniraptora is already everything closer to
Neornithes than to Ornithomimus. Enigmosaurus has no place in its
definition (wasn't even considered theropod by Gauthier when he named the
clade...).
|--"Enigmosauria" Keesey,
unofficial
Hmm. Tom Holtz HAS used that term on the
list. At last, I can stop writing "the segnosaur-oviraptorosaur group"
when on a public forum!
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|--Elmisauridae
Caenagnathidae is the correct name.
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|--Segnosaurinae
(?=Enigmosaurinae)
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|--Enigmosaurus
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|--Erlikosaurus
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`--Segnosaurus
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`--Therizinosaurinae
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|--Nanshiungosaurus
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`--Therizinosaurus
Segnosaurinae has priority over
Enigmosaurinae. Why oh why would you place Nanshiungosaurus and
Therizinosaurus together? There's nothing comparable between the two. The
former is known from vertebrae and a pelvis, the latter from fore and
hindlimbs. I suppose if the "Chilantaisaurus" zheziangensis ungual does
belong to the Nanshiungosaurus holotype, we could compare that, but I don't
think you did...
`--Eumaniraptora
|--+--Aves
==(1)
| | |?-Alvarezsauridae
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`---->Pygostylia
| `--Dromaeosauroidea
(implied Matthew & Brown, 1922)
| |--Archaeopterygidae
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|--Archaeopteryx
| | `--Rahonavinae
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|--Rahonavis
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`?-Vorona
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`---->Dromaeosauridae
Eumaniraptora is (Deinonychus + Neornithes) so
would be the node uniting Aves and Dromaeosauroidea in your phylogeny.
Vorona is really much more derived than Rahonavis, even having a few euornithine
characters. It's some type of ornithothoracine...
2) hyperextendable toe secondarily reduced and (?)lost in tyrannosaurs;
L-shaped cannon bone of the foot; and the T-shaped bone of the jugal region
(I can't remember what the specific bone is called but it was used to unite the
tyrannosaurs with the ornitholestians and the allosaurs)
The bone is the quadratojugal. It's also
dorsally expanded in Sinosauropteryx by the way.
Mickey Mortimer
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