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Re: giant birds
In a message dated 11/11/99 3:36:52 PM EST, k.clements@auckland.ac.nz writes:
<< I'm afraid I can't agree with George that "BCF and cladistic analysis
are not opposed; they work together. I use available dinosaur
cladograms to trace the evolution of avian features." I can't put it
any better than Luis Chiappe, who said (Nature 378: 349-355 1995):
"assumptions about evolutionary processes or adaptational scenarios,
such as these, are misleading when identifying historical
relationships. Phylogenetic reconstruction should be based exclusively
on the hierarchical distribution of homologies among taxa, and a given
phylogenetic hypothesis can only be rejected by providing an
alternative hypothesis for which supporting evidence outweighs that of
the original hypothesis." >>
Sorry, Chiappe is wrong. There are many ways to reject cladograms other than
by building bigger cladograms. You can check them against stratigraphy and
biogeography, and you can check them against functional anatomy. You can even
reject cladograms on the basis of common sense, if you like. There are still
too many holes in cladistic analysis itself for it to be granted this kind of
authoritarian philosophical position. Nobody has to accept any cladogram just
because its author can provide statistical arguments, for example, that show
how good it is.
But suppose for the moment that Chiappe is right, and that only by winning a
synapomorphy war--that is, only by building a cladogram from a larger, more
inclusive character matrix--can you reject a previous cladogram. This
paradigm will eventually render cladistics untestable, unfalsifiable, and
thus unscientific by producing the Final Cladogram, one that is done from the
largest possible character matrix by the best possible algorithm. This
cladogram--by construction--cannot be tested against other cladograms,
because it already includes all the information from which the other
cladograms could be produced. Since Chiappe admits no >other< way to test the
Final Cladogram, it is untestable, and therefore unscientific. Too bad,
that's the way it works. We haven't reached the Final Cladogram in
dinosaurology yet (although the way Tom Holtz is scavenging characters these
days, we soon will :-), but if we do, how will we know it really is the True
Phylogeny? Or do we enter the fool's Paradise and simply >declare< it to be
just that?
You still haven't figured out BCF and its relationship to dinosaur
cladistics. I did not produce a phylogeny from the idea that dinosaurs
originated as arboreal archosaurs. I >arrived at< this idea from examining
numerous published phylogenies and then extrapolating the likely features of
the various common ancestral forms from them. Phylogeny leads to BCF, not the
other way around.