phylogenies from Sereno 1999, pt. 1

["Thomas R. Holtz, Jr." <th81@umail.umd.edu>]

Hi, everybody!

Have been working my way through the details of the Sereno 1999 (Science
284: 2137-2147) phylogenies, and thought some people might be interested.
There are actually more taxa involved in these phylogenies than shown in the
resulting cladograms (details were condensed for the figures).

The analyses are broken up into eight different studies, then the results
were combined for the figures.  Here are the different analyses (taxa with
"*" were operational taxonomic units in that study):

I. Dinosauria
Ancestor *
_Marasuchus_ *
Dinosauria
  Ornithischia
    _Pisanosaurus_ *
    unnamed
      _Lesothosaurus_ *
      Genasauria
        Thyreophora *
        Neornithischia
          Ornithopoda *
          Marginocephalia *
  Saurischia
    Sauropodomorpha
      Prosauropoda *
      Sauropoda *
    Theropoda
      _Eoraptor_ *
      unnamed
        Herrerasauridae
          _Herrerasaurus_ *
          unnamed
            _Chindesaurus_ *
            _Staurikosaurus_ *
        Neotheropoda
          Ceratosauria *
          Tetanurae *

II. Thyreophora
_Lesothosaurus_ *
Genasauria
  Neornithischia *
  Thyreophora
    _Scutelloaurus_ *
    unnamed
      _Emausaurus_ *
      unnamed
        _Scelidosaurus_ *
        Eurypoda
          Stegosauria
            _Huayangosaurus_ *
            Stegosauridae
              _Dacentrurus_ *
              Stegosaurinae
                _Tuojiangosaurus_ *
                _Lexovisaurus_ *
                Other stegosaurines *
[note: haven't run the matrix, and am not certain how the last three OTUs
resolve]
          Ankylosauria
            Nodosauridae
              _Hylaeosaurus_ *
              Nodosaurinae
                _Pawpawsaurus_ *
                Other nodosaurines *
            Ankylosauridae
              _Gargoyleosaurus_ *
              unnamed
                _Minmi_ *
                _Shamosaurus_ *
                Ankylosaurinae
                  _Gastonia_ * 
                  _Pinacosaurus_ *
                  Other ankylosaurines *
[again, uncertain of details from _Shamosaurus_ on up]

III. Ornithopoda
Thyreophora *
Neornithischia
  Marginocephalia *
  Ornithopoda
    Heterodontosauridae *
    Euornithopoda
      Hypsilophodontidae * (yeah, I know: monophyly of this group is not
tested here)
      Iguanodontia
        _Muttaburrasaurus_ *
        _Tenontosaurus_ *
        Dryomorpha
          Dryosauridae *
          Ankylopollexia
            Camptosauridae *
            Styracosterna
              _Probactrosaurus_ *
              Hadrosauriformes
                Iguanodontidae *
                Hadrosauroidea
                  _Ouranosaurus_ *
                  unnamed
                    _Protohadros_ *
                    Hadrosauridae
                      Hadrosaurinae *
                      Lambeosaurinae *

IV. Marginocephalia
Thyreophora *
Neornithischia
  Ornithopoda *
  Marginocephalia
    Pachycephalosauria
      _Stenopelix_ *
      unnamed
        _Goyocephale_ *
        unnamed
          _Homalocephale_ *     
          Pachycephalosauridae
            _Stegoceras_ *
            Pachycephalosaurinae
              unnamed (Asian clade)
                _Tylocephale_ *
                _Prenocephale_ *
              unnamed (Lancian North Am. clade)
                _Stygimoloch_ *
                _Pachycephalosaurus_ *
    Ceratopsia
      _Psittacosaurus_ *
      Neoceratopsia
        _Chaoyangsaurus_ * (with a ref.: S. Zhao, Z. Cheng & X. Xu, in
press, JVP!!)
        unnamed
          _Archaeoceratops_ *
          unnamed
            _Leptoceratops_ *
            Coronosauria
              Protoceratopsidae *
              Ceratopsoidea
                _Montanoceratops_ *
                unnamed
                  _Turanoceratops_ *
                  Ceratopsidae
                    Ceratopsinae *
                    Centrosaurinae *


To be concluded...

                        Thomas R. Holtz, Jr.
                        Vertebrate Paleontologist
Deptartment of Geology                  Director, Earth, Life & Time Program
University of Maryland                  College Park Scholars
College Park, MD  20742       
Webpage: http://www.geol.umd.edu        Phone:301-405-4084
Email:tholtz@geol.umd.edu               Fax:  301-314-9661