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pterosaurian monophyly
I forwarded to Kevin Padian one of David Peters' (DPTEROSAUR@AOL.COM)
recent messages about pterosaur systematics. Kevin had already heard
about it from someone else on the list (I'm guessing Jennifer
Allen)... In any case, Kevin had a few comments that he said I could
share:
Pterodactyloids may or may not be monophyletic (I think they are,
based on current analyses), but there is a methodological question
here that ought to be of central importance to readers of this list.
One of the tenets of phylogenetic systematics is that you can't
assert convergence, you have to demonstrate it by showing that
another phylogeny is better supported. And that means a full
cladistic analysis with all the characters. This argument also
applies to David Peters's statements (from some weeks ago) that the
hindlimb characters of pterosaurs and dinosaurs that are interpreted
as correlated with bipedalism (by me, at least, but what do I know?)
are probably convergent. First, you have to show that pterosaurs
belong to another group -- again, by a full cladistic analysis,
demonstrating that those of Gauthier, Sereno, and me are not as well
supported as another explicit hypothesis. This would require a lot
of other characters, not just statements that the characters we used
are probably convergent and therefore can be disregarded. The
phylogenies that are currently accepted may well be problematic or
completely wrong, especially if I had anything to do with them.
However, the question is one of method, and we don't want to confuse
ideas of how the evolutionary process worked with ideas of how that
pattern is structured; instead, we want to test them against each
other. For details on this, people can read any recent issues of
Systematic Biology or Cladistics, or peruse some of the textbooks on
phylogenetic systematics such as those of Wiley, Brooks & McClennan,
Cracraft and Eldredge, and so on.
-- kp
--
Mickey Rowe (rowe@lepomis.psych.upenn.edu)